I would like to acknowledge the contribution of several individuals who were of immeasurable help in the preparation of this paper. Marcy Burchfield of the Neptis Foundation and Peter Gozdyra of the University of Toronto Cartography Department produced the figures and Rosalind Chaundy of Gartner Lee compiled the greenlands definitions in Appendix A. Two of my fellow ecologists at Gartner Lee, Dale Leadbeater and Brian Henshaw, acted as a sounding board for many of the ideas expressed in the paper and provided valuable comments on an earlier draft. My thanks also go to Tony Coombes (Neptis), Richard White (Neptis), and Pamela Blais (Metropole Consultants) for their advice and guidance.